A. L. V.  Davis, ?Dung beetle diversity in South Africa: influential factors, conservation status, data inadequacies and survey design?, African Entomology, vol. 10, pp. 53-65, 2002.

G.  Halffter, Favila, M. E., and Halffter, V., ?A comparative study of the structure of the scarab guild in Mexican tropical rain forests and derived ecosystems?, Folia Entomológica Mexicana, vol. 84, pp. 131-156, 1992.

M. A.  McGeoch, Van Rensburg, B. J., and Botes, A., ?The verification and application of bioindicators: a case study of dung beetles in a savanna ecosystem?, Journal of Applied Ecology, vol. 39, pp. 661-672, 2002.