A. L. V.  Davis, ?Biogeographical groups in a southern African, winter rainfall, dung beetle assemblage (Coleoptera: Scarabaeidae) - consequences of climatic history and habitat fragmentation?, African Journal of Ecology, vol. 31, pp. 306-327, 1993.

A. L. V.  Davis and Dewhurst, C. F., ?Climatic and biogeographical associations of Kenyan and northern Tanzanian dung beetles (Coleoptera: Scarabaeidae)?, African Journal of Ecology, vol. 31, pp. 290-305, 1993.