A. L. V.  Davis, ?Dung beetle diversity in South Africa: influential factors, conservation status, data inadequacies and survey design?, African Entomology, vol. 10, pp. 53-65, 2002.

A. L. V.  Davis, Scholtz, C. H., and Deschodt, C., ?A dung beetle survey of selected Gauteng nature reserves: implications for conservation of the provincial scarabaeine fauna?, African Entomology, vol. 13, pp. 1-16, 2005.