| Publication Type: | Journal Article |
| Year of Publication: | 2002 |
| Authors: | S. Villalba, Lobo, J. M., Martin-Piera, F., Zardoya, R. |
| Journal: | Journal of Molecular Evolution |
| Volume: | 55 |
| Pagination: | 116-126 |
| Keywords: | iberian beetles, nesting behavior, onthophagini, Scarabaeinae |